I think Jon is right, from the Gigabyte manual on the Z390 Designare:
1 x PCI Express x16 slot, running at x16 (PCIEX16)
* For optimum performance, if only one PCI Express graphics card is to be installed, be sure to install it in the PCIEX16 slot.
1 x PCI Express x16 slot, running at x8 (PCIEX8)
* The PCIEX8 slot shares bandwidth with the PCIEX16 slot. When the PCIEX8 slot is populated, the PCIEX16 slot operates at up to x8 mode.
1 x PCI Express x16 slot, running at x4 (PCIEX4)
2 x PCI Express x1 slots
(All of the PCI Express slots conform to PCI Express 3.0 standard.)
1 x M.2 Socket 1 connector for an Intel® CNVi wireless module (CNVI)
The other question is, will it make any difference? If you are keeping the 1070 in your sidebar, probably not given a GTX1080 at 8 lanes is near enough to make no difference to 16 lanes (
https://www.techpowerup.com/reviews/...press_Scaling/), and even 4 lanes is within spitting distance (one of the final charts shows x4 has 96% of the performance of the 16x).
edit: Was too slow, you already got your answer.