Video on the D5000 leaves a lot to be desired, I have it. It may come in use if you really need it, but considering it doesn't even have autofocus, unless you like constantly manually focusing during filming, you're better off getting a camcorder instead.
You mention you'd like to photograph animals - what types, because you'll most likely need some long lens? Something to consider maybe that the canon 300mm f4 has IS whereas the Nikon doesn't. Coupled with the 1.4tc you effectively have 630mm on a cropped sensor camera.
On the other hand you mention macro, and depending how serious you are, I *think* the Nikon 105mm is considered the best macro (anybody feel free to correct me if I'm wrong, I'm no expert).
Not really useful in helping you to make a decision I guess!